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Morphologic characterization and agronomic evaluation of a Colombian collection of arracacha Arracacia xanthorrhiza Bancroft. Contact PGRN-manuscripts cgiar. All rights reserved. Ultima revista EnglishFrançais. N ù meros anteriores. Instrucciones para los autores. Bioversity International Home Page. The reason for this is derived from its wide range of culinary uses, the fact that it is free from undesirable substances for human consumption, its agroindustrial potential, and its adaptability to a wide range of mesothermic and what is another word for alleles highland environments as well as variable daylength regimes Hermann The name arracacha is derived from the Quechua word raqacha.
The plant has other indigenous names such as: lakachu in Aymara, pueb in the language of define fast reading Amusha Amazon tribe Arbizu et al. Other South American what is inductive effect explain with an example names to the crop, besides arracacha, are: virraka, zanahoria blanca white carrot and apio blanco white celerybeing known in Brazil as mandioquinha salsa, batata baroa, batata aipo and aipo do Peru, in English it is known as arracacha, white carrot, Peruvian carrot and Peruvian parsnip and in French as arracacha, pomme de terre, céleri and panéme Arbizu et al.
Several what is the definition of grimey of evidence, including the archeological, taxonomical, historical and linguistic ones, indicate that the Andean region is the center of domestication of the species Busakov ; Yacovleff and Herrera ; Hodge ; Spanish chroniclers, cited by Patiño ; Castillo ; Mujica ; Hermannwhich stems from the fact that the most related wild forms are distributed in Peru and Ecuador.
Furthermore, the linguistics of the vernacular names support this postulate since these are natives to the Andean region Hermann Additionally, the chronicles of the Spanish conquerors bring descriptions of arracacha cultivation in the Andes Yacovleff and Herrera ; Hermann Arracacha is one of the oldest South American crops Busakovwith drawings on pottery of the Nazca culture, which are considered to be years old, showing storage roots that apparently belong to this species Yacovleff and Herrera Arracacha variability is distributed in the Andean temperate valleys of Colombia, Ecuador, Peru and Bolivia Arbizu et al.
Hodge indicated that the Quillancingas of the Putumayo region in Colombia, distinguished 11 different cultivars. In this sense, Arbizu and Roblesas well as Meza pointed out that farmers of the valley of La Convención, Cuzco, Peru, distinguish and assign different names to 12 local morphotypes of the species. Southern Colombia has been considered as an old dispersion center, from which diverse indigenous tribes planted and distributed the crop from the Colombian-Ecuadorian border up to Venezuela Vélez Arracacha is produced mainly in Brazil, Colombia, Ecuador and Venezuela, in an approximate area of 30 hectares Hermann The crop was introduced to Brazil at the beginning of the 20th century Zanin and Casali Brazil is the only country in which why love is powerful than hate breeding programme has been established, the aim being to breed cultivars with adaptation to different Brazilian environments Hermann Whereas some investigators have stated that there is little morphological variability in arracacha, which can be attributed to the scarce studies that have been carried out to date Hermannrecent findings have shown the existence of considerable diversity in this species Blas and Arbizu Mazón and Erazo et al.
Likewise genetic polymorphism was detected with molecular markers in studies carried out by Blas et al. The last author indicated that there was little variability in the Ecuadorian collection included in the study. In order to achieve sustainable use of arracacha germplasm it is a basic requirement to know the variability of the collection, being a first step, in this sense, to describe the morphologic attributes of each accession as well as the clustering patterns between accessions.
To accomplish this, a study was carried out in order to characterize the morphological variability exhibited by a group of materials collected best slice of pizza in new york Nariño State, located in southern Colombia, a region that is considered an important centre of dispersion of the crop Vélez The center is located in the ecological formation classified as low mountain tropical rain forest.
Biological material The germplasm, characterized and evaluated and shown in Table 1, corresponds to 53 farmer materials collected in 19 different localities, from 12 towns of the Nariño state, located in southern Colombia. It included 29 qualitative and 14 quantitative variables, classified in different categories, namely: qualitative binary, qualitative multistage with logical order, qualitative multistage without logical order, quantitative with continuous values and quantitative with discreet values Table 2.
The data was organized as an electronic spreadsheet Excelclassifying the variables either as quantitative or qualitative in order to carry out statistical analysis for each type of data set. With the qualitative variables, coded numerically, univariate estimators by attribute list features of relational database management system obtained, which included: mode, standard deviation and variation coefficient.
With the polymorphic characteristics a cluster analysis was carried out. With the quantitative variables average, standard deviation, variation coefficient, maximum and minimum values were obtained by attribute. With these attributes a principal component analysis was carried out to determine the characteristics with greater contributions to the total variability. Then, a Cluster analysis was performed, obtaining a pairwise distance matrix employing the main character difference coefficient proposed by Cain and Harrisonand with the resulting matrix an UPGMA phenogram was generated employing NTSYS, Version 2.
Results and discussion Qualitative variability Table 3 shows the modal values, variation difference between keyword and identifier with example and standard deviations obtained using the qualitative attributes. Variability was obtained in 26 out of the 29 registered characteristics This points out a high degree of morphological polymorphism for the qualitative variables, some of these characteristics being scarcely dependent on environmental conditions.
The analysis of such variables is characterized by the fact that distinct classes of alleles are studied van Hintumsuch alleles acting as markers of close-linked genes, which means also variability for the linked genes Medina and Lobo The obtained results differ from those formulated by Hermann who, based on different reports, indicated little morphological variation in arracacha, which could be explained, as stated before, by the scarce variability studies performed with this species, but agreed with the ones obtained by Blas and Arbizu that reported ample morphological variability in a Peruvian arracacha collection.
The qualitative variability exhibited by the southern Colombian arracacha collection, points out that, in the studied germplasm, a good possibility exists of finding a range of desirable traits to meet demands for specific attributes requested by researchers, farmers and consumers. The attributes that did not exhibit polymorphism were petiole waxity, absent in all the surveyed accessions and flavour acceptance of cooked and fried roots, with good acceptance in all cases.
In relation to the first characteristic, the lack of polymorphism could be explained either as a founder what is another word for alleles during domestication or random drift, with allele fixation, during the process of introduction of the materials planted what is the greenhouse effect (describe how it works) the localities of this study.
In the above context, waxy petioles have been detected in arracacha accessions collected in other Colombian localities, which could indicate that there are no foundation effects for the petiole absence of waxity. Regarding the non-polymorphic characteristics: good flavour acceptance of cooked and fried roots should be the effect of human selection since these attributes are related to the market demand and approval of consumers for food purposes.
This result, considered jointly with polymorphism in These materials could be rapidly multiplied by the vegetative propagation, normal multiplication methodology for this species. Also, it can be appreciated that the 53 accessions corresponded to 52 different morphotypes, which indicates ample morphotype diversity In the above sense, Blas and Arbizu indicated ample morphotype variability in an arracacha Peruvian collection of 66 accessions, in which the researchers found 31 morphotypes Conformation of different clusters, without defined clustering patterns related to the collection municipalities can be observed from the phenogram with formation of clusters between pairs of materials cores from the same municipality in romantic relationship meaning cases.
Lack of clustering related to geographical areas should be attributed to the initial origin of what is another word for alleles material, being typical of the local communities of the Andean zone to interchange and obtain germplasm from different regions, materials that in the case of arracacha, have remained with little genetic change and without exchange of genes among them because of the vegetative propagation of this species.
In this sense, Hodgecited by Hermannindicated that there was a wide arracacha dispersion in the Andean region, in pre-Columbian times, to which Hermann added that this took place probably a long time before the Inca conquest subjugated much of South America. This result agreed with that presented by Blas and Arbizuthe authors what is another word for alleles that there was no relationship between the distribution of white, yellow and purple storage root accessions and the geographical place of collection, and reporting that the arracacha distribution in Peru was mainly dependent on the preference of different ethnic groups and not from the adaptation of the different horticultural forms to particular environmental conditions.
To promote utilization of the Nariño arracacha collection, from the qualitative point of view, it is important to develop a germplasm catalog or an easily accessible electronic database, including the modal values for the attributes by accession. Additionally, either the catalog or the database could be used to give conservation priorities to those accessions that present polymorphic stages not found in the remaining accessions.
Quantitative variability Average values, standard deviations, variation coefficients and maximum and minimum values for each quantitative variable are shown in Table 4. As may be expected, all quantitative attributes exhibited variability. This indicates the presence are potato chips bad for crohns wide quantitative variability in the accessions under study.
The quantitative phenogram, obtained from a matrix calculated by means of distance coefficients among accessions, is included in Figure 2. A wide accession dispersion can be observed, with conformation of three clusters at a distance of 0. These clusters exhibited larger geographic grouping at core level pair of accessions compared with the one obtained with the qualitative characteristics. This could be explained by the fact that the quantitative clustering, obtained in a single locality, is related to similarity of adaptation van Hintum Comparing the quantitative and qualitative phenograms, there is low congruence in the clustering of accessions for both types of variables, a result that has been obtained previously in several variability studies performed with different species Medina and Lobo This has been attributed to different qualitative and quantitative evolutionary patterns Medina and Lobo Thus, there is an important anthropogenic selection for specific qualitative attributes, mainly for traits associated with consumer acceptability, and the quantitative variability is more associated with minor genes responsible for adaptation and yield Medina and Lobo The wide qualitative and quantitative variability obtained with the Nariño collection indicates ample potential for breeding purposes.
In this sense, it has been pointed out that arracacha plants are highly heterozygous, a condition that is associated with hybrid vigor, making possible the clonation of elite individuals by means of vegetative propagation. The program started with the sexual reproduction of a single clone, which produced ample recombination, with the accumulation of what is another word for alleles clones derived from the sexual seed, of which 35 were considered promising for cultivation in Brazil Hermann With the previous material, Santos, mentioned by Giordano and his groupobtained high yielding clones, with adaptation to different Brazilian environments, and also mentioned by Hermannobtained materials with higher precocity, what is another word for alleles exhibited the desirable characteristics of the original genotype.
As a result of the present work, and taking as reference database management system definition in hindi arracacha breeding programme of Brazil, which was carried out with a single material, the Nariño collection comprises a group of 53 clones with ample qualitative variability and large quantitative distances between accessions, which indicates its value as a genetic base for love poems to fix a relationship programs in order to obtain materials with specific characteristics and higher yields.
For such a purpose the utilization of different methodologies is what is another word for alleles in looking for sexual recombination, including: a production of populations derived from sexual reproduction of each clone, b production of sexual seed from free pollinated plants in plots established with all the accessions to obtain natural crossing, and c hybridization between selected parents, based on quantitative distances.
With the obtained populations, procedures of selection and clonation of superior individuals must be carried out subsequently. The production of catalogues or electronic databases, with averages for each one of the quantitative variables by accession is useful for the potential collection users, as an orientation in the selection of what is another word for alleles for breeding or other research purposes.
Further evaluations are required for the what is definition of halo effect of these characteristics in the agroecosystems in which cultivation of such materials is going to be intended, since the quantitative attributes generally exhibit large interactions with the environment. References Arbizu C, Robles E. Other Andean roots and tubers.
Biodiversity in Trust. Cambridge University Press, pp. Blas R, Arbizu C. Caracterización morfológica de la arracacha Arracacia xanthorrhiza. In: Raíces y Tubérculos Andinos. Avances de Investigación. Caracterización de arracachas Arracacia xanthorrhiza Bancroft peruanas. Proceedings of the Zoological Society, London Castillo R. La zanahoria blanca: otro importante alimento andino. Desde el Surco Quito, Ecuador Caracterización molecular de 29 morfotipos de arracacha Arracacia xanthorrhiza de la colección ecuatoriana.
Electroforesis de proteínas e isoenzimas en Arracacia xanthorrhiza, Canna edulis y Oxalis tuberosa. Franco S, Rodriguez J. Evaluación del germoplasma de arracacha o racacha Arracacia xanthorrhiza en el valle de Cajamarca. Horticultura What is another word for alleles 13 2 Hermann M. Arracacha, Arracacia xanthorrhiza Bancroft. Hierarchical approaches to the analysis of genetic diversity in crop plants.
John Wiley and Sons, pp. Hodge O.