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Dirección para correspondencia. SUMMARY: Catfishes constitute a highly diversified, cosmopolitan group that represents about one third of all freshwater fishes and is one of the most diverse Vertebrate taxa. The detailed study of the Siluriformes what are some practical applications of phylogenetic analysis, thus, provide useful data, and illustrative examples, for broader discussions on general phylogeny and macroevolution.
In this short note I briefly expose how the study of this remarkably diverse group of fishes reveals an example of highly homoplasic, complex 'mosaic' morphological evolution. En esta comunicación expondré brevemente cómo el estudio de what are some practical applications of phylogenetic analysis grupo notoriamente diverso revela un ejemplo de amplia homoplasia y un complejo «mosaico» de evolución morfológica. The catfishes, or Siluriformes, found in North, Central and South America, Africa, Europe, Asia and Australia, with fossils inclusively found in Antarctica, constitute a highly diversified, cosmopolitan group, which, with more than species, represents about one third of all freshwater fishes and is one of the most diverse Vertebrate taxa e.
Burgess, ; Diogo, ; Teugels, The detailed study of the Siluriformes can, thus, provide useful data, and illustrative examples, for broader discussions on evolutionary biology Diogo, In this aspect, one of the points that most struck me in my research on these fishes in the last years is the rather high level of homoplasy and complexity of their morphological evolution.
Catfishes as an example of complex, mosaic morphological evolution. This subject, it should be noticed, is deeply related with my own personal scientific development. When I first started the observations and comparisons on catfishes, the impression was that a great part of the evolutionary changes concerning the major morphological systems of those fishes were somewhat 'oriented', in a somewhat 'simple' way.
Of course, I felt the incredible complexity and diversity of catfishes, a complexity and diversity surely resulting from several homoplasic events. This was precisely the main reason to choose this amazing group of fishes as a case study for discussing general what are some practical applications of phylogenetic analysis on phylogeny and macroevolution. But the exam of more and more morphological phylogenetic characters in numerous catfishes, reaching to a total of characters in 87 genera representing all the 32 extant families of the order, and specially the subsequent results obtained from the cladistic analysis of those characters Diogo, The strict-consensus cladogram obtained in that cladistic analysis, of which the main results are briefly summarised in Figure 1 showing the relationships between the extant catfish families, had a lenght of steps and a Consistency Index CI of 0.
That is. One could eventually argue that such a what are some practical applications of phylogenetic analysis of homoplasy could perhaps be exclusively related with an eventual incorrectness of the phylogenetic results obtained in the cladistic analysis of Diogo However, the. The CI value obtained in Diogo's work is significantly higher, for example, than the CI value obtained in the two other published cladistic studies on catfish higher-level phylogeny, which in fact included less phylogenetic characters than Diogo's work Mo,included characters and had a CI of 0.
Thus, if instead of the phylogenetic results of Diogo one would take the phylogenetic results of these two other cladistic studies, the level of homoplasy would even be greater. Also, a CI of 0. In reality, the study of catfishes effectively seems to provide an example of rather homoplasic, complex 'mosaic' morphological macroe volution. The rather complex and 'mosaic' morphological evolution of catfishes can be illustrated by a practical example concerning one of the best supported clades within the order: that formed by the Asian Sisoroidea Amblycipitidae, Akysidae, Sisoridae and Erethistidae and the South American Aspredinidae see Fig.
The first author providing evidence to suggest that the South American aspredinids were related to Asiatic taxa was Ferraris. Mo, in the first explicitly phylogenetic analysis of siluriform higher-level phylogeny, supported this view, placing the aspredinids as either basal to or in a polytomy with a clade containing the Asian amblycipitids, akysids and sisorids.
Somewhat similar hypotheses were suggested subsequently by de Pinnawho placed the Aspredinidae in a polytomy also including these three Asian groups, as well as the Amphiliidae from Africa and the Loricarioidea what does damaged skin mean in spanish South America. InChen provided further evidence to place the Aspredinidae as the sister group of a clade composed of the Asian families Amblycipitidae, Sisoridae and Akysidae.
This hypothesis was subsequently strongly supported by de Pinnawho considered that the Sisoridae of previous authors was a paraphyletic assemblage, with a subunit of it which he named Erethistidae: see Fig. This view was posteriorly corroborated in de Pinna's overview on the phylogenetic relationships of Neotropical catfishes. The cladistic analysis of the author of the present work Diogo, on siluriform higher-level phylogeny strongly supported the hypothesis of de Pinnawith the Neotropical Aspredinidae being grouped in a monophyletic clade together with the Asian Sisoridae and Erethistidae, as shown in Figure 1.
Thus, there is strong evidence accumulated in the last 15 years supporting a close relationship between the South American Aspredinidae and the Asian Sisoroidea families Amblycipitidae, Akysidae, Sisoridae and particularly Erethistidae see Fig. Attending to the strong evidence supporting the close relationship between the Aspredinidae and the Asian Sisoroidea families Amblycipitidae, Akysidae, Sisoridae and particularly Erethistidae, it is thus very interesting to notice that, as stressed by de Pinnathe aspredinids share some" striking" derived anatomical features with other why are predator prey relationships density dependent groups such as the Chacidae or the Doradidae see Fig.
The aspredinids and the chacids share, effectively, some rather peculiar, rare morphological characters that, attending to the strong evidence supporting the clade Sisoroidea Fig. In particular, the configuration of the posterodorsal region of the skull in the members of these two families is remarkably similar, as stressed Chardon For example, both these groups present a well-developed, deep fossa between the posttemporo-supracleithrum, the parieto-supraoccipital and, eventually, the epioccipital see Fig.
Such a fossa is only found, besides these groups, in the What are some practical applications of phylogenetic analysis erethistids see Fig. Also, both the Aspredinidae and Chacidae present a well-developed, dorsal lamina of the Weberian apparatus contacting with the dorsal surface of the body see Fig. Another peculiar, and also rather rare, feature present in the Aspredinidae and the Chacidae is the markedly thin and mesially extended dorsomesial limb of the posttemporo-supracleithrum see Fig.
But the morphological similarities between chacids and aspredinids are not only restricted to the configuration of the structures of the posterodorsal region of the cranium. For instance, in both these groups the prevomer is missing see Fig. The absence of the prevomer is a highly peculiar and rare character among the order Siluriformes, only occurring in a few other catfishes such as, for example, the members of the pimelodid genus Microglanis and of the scoloplacid genus Scoloplax.
According to Chardon, the anatomical similarity between the chacids and the aspredinids could probably what does dirty bird mean in text associated to a homoplasic adaptation to a peculiar 'burying' behaviour exhibited by the members what are some practical applications of phylogenetic analysis these two groups. But, as mentioned above, the rather "mosaic", complex combination of peculiar characters present in the Aspredinidae makes that these fishes also share some remarkable peculiar features with other non-Sisoroidea groups that are morphologically very different from the Chacidae, as, what is difference between variable and data type example, the Doradidae.
Some of these characters concern, for example: the prominent dorsolateral projections of laminar bone of the mesethmoid; the mesocoracoid arch and the main body of the scapulo-coracoid being undistinguished from each other; the presence of the highly developed anterior process of dorsal condyle of pectoral spine; and the presence of the well-developed anteroventral lamina of the preopercle see Fig.
These characters, and other features, what are some practical applications of phylogenetic analysis inclusively what are some practical applications of phylogenetic analysis Friel to propose a close relationship between the aspredinid and the doradoid catfishes i. However, as explained above, the phylogenetic results of Ferraris, Mo, de Pinna ,Chen, Diogo et al. In fact, the point that I would like to stress here is that, even if all those phylogenetic studies supporting the close relationship between the Aspredinidae and the Asian Sisoroidea see Fig.
The situation could be briefly what are some practical applications of phylogenetic analysis as follows. Let's take a 'triangle' with the Aspredinidae in the center and with 1 the Asian Sisoroidea family Erethistidae, 2 the Doradidae, and 3 the Chacidae in the extremities of the triangle, respectively see Fig. Thus, the remarkable peculiar similarities found between, in the one hand, the Aspredinidae and the Doradidae, and, in the other hand, the Aspredinidae and the Chacidae, are thus assigned to homoplasic events, thus revealing a particularly complex, 'mosaic' homoplasic evolution within these groups see Fig.
Therefore, the point is that the choice of any extremity of this 'triangle' would imply necessarily a series of homoplasic events between the Aspredinidae and the groups represented in the other two extremities. In reality, as explained above, the phylogenetic scenario proposed by the studies of Ferraris, Mo, de Pinna ,Chen, Diogo et al. Thus, the high level of homoplasy illustrated in this example cannot simply be explained by the choice of an erroneous phylogenetic scenario, but rather by a seemingly truly, highly complex 'mosaic' morphological evolution.
This example also illustrates an important point that has been stressed by authors such as e. As stated by Klassen et al. However, such a 'confidence measurement' only makes full sense when comparing levels of homoplasy exhibited by cladograms concerning a somewhat similar number of characters referring to a similar type of data set in a same biological group. Indeed, there is no reason to think that all different biological groups, even of a relatively similar size i.
If one obtains a certain consistency index A in the cladogram of a group X and this index A is smaller than that of a cladogram B concerning a group Y, there is no reason to consider, a priori, even if the number of taxa and characters analysed in both cases is somewhat similar, that the cladogram B is 'probably more do dating websites actually work than the cladogram A. So, as stated in what are some practical applications of phylogenetic analysis premises of the cladistic paradigm, but sometimes confused in practice, a cladogram with a relatively small consistency index could eventually be simply related with the occurrence of a truly high level of homoplasy in the group to which the cladogram refers, and not necessarily to a 'bad cladogram'.
The interpretation of the information given in a cladistic analysis by homoplasy indexes such as the consistency index should thus be made with much caution, case by case, taking into account not only the number of characters and of terminal taxa included in the cladistic analysis, but also the type of characters and the biological group to which the cladistic analysis refers.
Cambray Albany Museum of Grahams to wnG. Laleyé Université Nationale du BeninR. What is the meaning of literally in tamil, J. Williams and S. Duhamel Museum National D'Histoire Naturelle for kindly providing a large part of the biological material used in my studies of catfishes during the last years. I would also like to acknowledge I.
Doadrio, I. Peng, G. Teugels, R. Vari, S. Weitzman, T. Abreu, A. Zanata, B. Kapoor, F. Meunier, S. He, D. Adriaens, F. Wagemans, C. Oliveira, E. Parmentier, and specially M. Skelton,M J. Stiassny, F. Poyato-Ariza, G. Arratia, T. Grande, M. Ebach, A. Wyss, J. Waters, B. Perez-MorenoG. Cuny, A. Choudhury, M. Vences, S. Weitzman, L.
Cavin, F. Santini, J. Briggs, L. Gahagan, Philiphe J. Maisey, M.
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