realmente extraГ±amente
Sobre nosotros
Group social work what does degree bs stand for how to take off mascara with eyelash extensions how much is heel balm what fifference myth mean in old english ox power bank 20000mah price in bangladesh life goes on lyrics quotes full form of cnf in export i love you to the moon and back meaning difference between blood and lymph composition punjabi what pokemon cards are the best to buy black seeds arabic translation.
Correspondence to :. It was found that the forestomach of yak consisted of the following three parts, rumen, reticulum and omasum, which were composed of the mucosa, submucosa, muscularis and serosa. In addition, the mucosal epithelium was covered with stratified squamous epithelium, with part of keratinized the shallow cells.
Rumen, the mucosa of which formed ligulate papillae varying in size and shape, was no muscularis mucosa. Reticulum, consisted of a surface epithelium that invaginated to various extent into the lamina propria, formed various folds in shape, namely, grid-like small rooms. Furthermore, there are many secondary folds densely covered with keratinized papillae.
The most striking feature of the omasum was to be formed the laminae omasi varying in length, with short and rough papillae distributing on both sides. Taken together, there was no glands within the mucosa and lamina propria of forestomach of yak, where diffuse lymphoid tissues can be observed clearly. It is, therefore, believed that the yak forestomach may have evolved those specific structural characteristics in response to the unique living environment and dietary habits impose on the Qinghai-Tibetan Plateau.
Se encontró que el preestómago del yak constaba de tres partes: rumen, retículo y omaso, compuestas de mucosa, submucosa, muscular y serosa. En el rumen, la mucosa formó papilas linguladas why nutrition food is important variaron en tamaño y bllood.
The yak Bos grunniensas a year-round grazing animal, is a key species in the world, living in the Qinghai-Tibetan Plateau. They mainly live in alpine grassland in northwestern and southwestern China. Yaks may have acquired many special abilities and attributes to adapt harsh natural environment characterized by high altitude 3, m above sea levelvery low annual average temperature, short growing season and great seasonal variation in feed supply.
Ruminant's stomach has developed four separate compartments, each with its own morphological particularities. The first three parts are rumen, reticulum and omasum, commonly known as forestomach Clauss et al. Difference between blood and lymph composition there has been considerable researches into the organization of the stomach in cattle Vivo et al. In this study, we observed morphological structure of the forestomach through method of gross anatomy, and then lymp histological structure by general staining and special histochemical method, to understand the morphological and physiological function of the yak forestomach better, especially adaption for the balance of whole-body energy homeostasis in short growing season, and to provide a resource for research into metabolism copmosition enable yaks to survive extreme environments in the Qinghai-Tibetan Plateau, China.
All research protocols used in the current experiment were approved by Animal Ethics Committee of the Qinghai province, China. Sampling and methods. Surface enlargement factor of rumen mucosa SEF was evaluated in this paper. There were many papillae in mucosa wall in lymmph, thus expanding absorption area of rumen. The size and density of papillae is not only related with animal species, but also with the availability of food resources and quality, therefore this method can be used to predict and evaluate the nutritional status of ruminants.
For laboratory analysis, the dehydrated specimens 1x2 cm were taken, then counted the density of papillae under dissecting microscope, and randomly measured the difference between blood and lymph composition and width of 20 papillae be accurated 0. Once the forestomach had been separated, small pieces of tissue were dissected for analysis. Gross anatomy structure. The stomach of the yak consisted of the following four parts, rumen, reticulum, omasum and abomasum.
The first three parts were commonly known as forestomach, which was focused on in this paper. There were sulcus cranials and sulcus caudalis in the front and rear end and dfiference ruminis sinster and sulcus ruminis dexter in the left difference between blood and lymph composition right side. The backpack and ventral sac around left and right longitudinal groove of the whole rumen were observed. The interface between rumen and reticulum was about 13 cm in diameter, in which was the vestibular of rumen in shape of pleats, it was the anterior column behind the vestibular.
Most of mucosa covered with dense papillae at the surface, especially abdominal sac, was brown or black. But the edge of adductor muscle was pale, and the mucosa of backpacks sac was gray. The papillae, which gathered into a carpet or brush-like, varied greatly in size and shape, such as flat, ligulate and leafy and so on.
The average length and density of papillae was about 0. However, the mucosa of adductor muscle and backpack sac that absented of papillae was smooth, then the sidewall of backpack sac only had low and sparse granular papillae. Reticulum was located in the forefront of rumen. There were great many grids varying in size and shape formed by uplift folds of mucosa, those folds interconnected each other and formed four, five-sided or hexagonal grids, which were about 1.
In addition, many lower secondary folds at the bottom of rooms did the second partition to those grids and formed small irregular grids, at the bottom of which densely covered with horny papillae, as well as on the folds. Moreover, the closer to the edge of pleats between rumen and reticulum, the smaller the rooms side effect of meaning in urdu, and bteween disappearing.
Furthermore, the lip of sulcus reticuli was looked upon as the diffefence of the bottom edge of opening and side edge of ostium reticulo-omasicum. Sulcus reticuli, also known as sulcus esophageus, started from the cardia, and downed along the vestibular of rumen and the right side of reticulum wall and connected with omasum ditch when reaching ostium reticulo-omasicum. The total length was about 23 cm. Behween mucosal folds on both sides of ditch, which was rich in muscle tissue difgerence so called the lip, was more smooth and slightly helical twist.
The omasum, lying mainly in the middle right side of the abdominal cavity, was oval in shape and had a mean capacity of 4. The oblique wall of omasum located in right front of diaphragmatic liver. Meanwhile, the facies visceralis was contact with reticulum, rumen and abomasum. Fundus of omasum was located in the lesser curvature toward right rear of omasum's bend, and the upper and lower ends of lesser curvature leaded to reticulum and abomasum, respectively.
Furthermore, the sulcus omasi was extending along the cavity surface of lesser curvature, where liquid and small particles feed would directly flow into abomasum from reticulum. Yak's sulcus omasi, which started at reticuloomasal orifice and ended at ostium omasoabomasicum, was approximately 4. Only some small folds and papillae at bottom of sulcus omasi can be seen obviously, but without laminae lympph. Interestingly, omasum's mucosa in yak was formed over hundreds of laminae omasi that was shaped like a crescent, attaching to the greater curvature of omasum wall and the free edge toward the lesser curvature.
Laminae omasi, which neatly and alternately arranged in a regular and divided omasum cavity into many narrow and ordered gap, were classified into 18 pieces of large, 10 cm wide, 20 pieces of medium, 5 cm wide, 40 pieces of small, 1 cm wide, and 60 pieces of linear gaps, 1 mm wide according to their width. According to anatomical features, the yak's reticuloomasal orifice was significantly bolod than omasoabomasal opening in diameter at winter, approximately 5 cm and 6.
What's the most important is that both of them varied greatly in diameter what is partnership in public health the different seasons, for example, approximately 6. Surface expansion coefficient of rumen. In yak, the surface expansion coefficient SEF of papillae in rumen mucosa was on average 5.
The degree of size and density of papillae varied considerably in different regions, so the lynph of SEF was relatively large difference. For example, on the dorsal wall, even though the degree of density was not same in different individuals, the size of papillae was smaller in general, corresponding to the smallest SEF compared to other regions, was only 2.
Comparing with the dorsal wall, however, the thick and full papillae were densely distributed on ventral wall, and the SEF was 9. Yet the SEF was greater than former, about 4. Histology structure. The histology structure of the yak's four stomachs was basically same under the light microscopy, that is, following by mucosa, submucosa, muscularis and serosa.
But the mucosa of forestomach belonged to non-glandular mucosa layer, only abomasum's mucosa layer had the glands. Rumen cardia. The mucosa of the cardia in the rumen formed tiny and low papillae distributed sparsely, and was consisted of partly keratinized stratified squamous difference between blood and lymph composition, which was composed of multi-layer cells varying in shape from upper to lower. A layer of basal cells near the basal membrane was low columnar, there were several layers of polygonal cells above basal layer, and then the flat cells, but the flat cells were partly keratinized on the outermost layer, where the cells with disappearing nuclei were dead, filling with keratin in the cytoplasm.
As for shallow corneum, was yet homogeneous and the surface cells are constantly shedding Fig. Deep cells, forming downwards a great many apophysis, were larger and went deep into lamina propria. Lamina propria was mainly composed of loose connective tissue without glands, containing collagen fibers. Due largely to the submucosa composing of a thin layer of loose connective tissue as well, so there was no clear boundary between lamina propria and compoosition, where some plasma cells, macrophages, fibroblasts and mast cells can be observed Fig.
There were lymphatics and abundant veins and capillaries, how does a normal relationship progression fewer arteries in smaller diameter in the loose connective tissue. Some large lymphocytes, which were easily stained by hematoxylin, were scattered in peripheral veins Fig.
The muscularis was difference between blood and lymph composition thicker and included two layers of smooth muscle, the inner oblique muscle and the outer longitudinal muscle Fig. The outer ring was about 1, While the inner oblique muscle was only 1, Veins and nerve plexus can be seen in connective tissue among muscle bundles.
The muscle cells, cytoplasms of which were stained lightly, were slender fibrous in shape, with oblate nuclei located below the basal membrane. In addition, each muscle cell was wrapped by the endomysium Fig. The adventitia was composed of connective tissue. It was rich in collagen and elastic fibers, in which there were a great many difference between blood and lymph composition vessels with the coarser diameter Fig. For vein, the wall was collapsed and its flat difference between blood and lymph composition was also irregular in shape.
For artery, the three-tier betwewn of wall was obvious. In addition, a large number of capillaries and lymph capillaries can be exhibited in the adventitia. There were abundant adipose tissue composed of a large number of fat cells near the muscularis. The nerve plexus, lymphocytes and fibroblasts distributed among connective tissue. What's the worth noting is that fibroblasts, which attached to collagen fibers with large nuclei stained lightly, were flat and its compposition belonged to basophilic.
Rumen vestibule. The surface of mucosa compodition covered with large and dense papillae, which were shaped like leaves, about 6, The mucosal epithelium was superficial keratinized stratified squamous epithelium. For the superficial compositon layer, the shallow layer was composed of several layers of flat cells without nucleus, while the nucleus of cells in deep layer were flat spindle.
The middle layer was composed of polygonal cells with round or oval nucleus. The basal layer was composed of low columnar cells with oval nucleus, which tightly packed and stained into adn. The deep epithelium com;osition small protrusions extending to the lamina propria. The lamina propria was composed of loose connective tissue without glands, and stretched papillae along the lympu epithelium to form a continuous strip.
Comparing with cardia, the lamina propria was relatively thin, containing small arteries, veins and capillaries Fig. Fibroblasts attached to collagen fibers, irregular fusiform in shape, with blue-purple cytoplasm and oblate nucleus Fig. The cytoplasm of plasma cells were abundant and were easily stained by hematoxylin.
Interestingly, there was no muscularis mucosa.
realmente extraГ±amente
Es la informaciГіn de valor
maravillosamente, es la pieza muy de valor
Bravo, que la frase necesaria..., la idea brillante
Bravo, son Гєtil su opiniГіn
No sois derecho. Soy seguro. Puedo demostrarlo. Escriban en PM, hablaremos.